Pugs were the official dog of the House of Orange in the Netherlands in the 16th century.
The word "pug" may have come from the marmoset monkey, which had a similar squashed face.
Pugs descended from short-muzzled wild dogs that lived in Asia over 2,000 years ago.
The distinctive wrinkles on a pug's face help channel scent molecules to their nose.
Pugs shed year-round and seasonally with two heavy shedding periods annually.
Brachycephalic pugs can suffer breathing issues, especially in heat and humidity.
Pugs' short legs and flat faces make them slow, inefficient runners.